The movies take Mattel's Hot Wheels toy cars and puts them in a ReBoot-esque situation, with racing somewhat reminiscent of Tron. This is a sequel series to Hot Wheels: World Race. A soundtrack was distributed by Sony BMG and toys based on the movies were sold in stores and given away in McDonald's Happy Meals.
The four movies constitute one continuous story and also continue from where World Race left off. The four movies that make up the series are:
- Hot Wheels: AcceleRacers – Ignition
- Hot Wheels: AcceleRacers – The Speed of Silence
- Hot Wheels: AcceleRacers – Breaking Point
- Hot Wheels: AcceleRacers – The Ultimate Race
Two years after the end of the World Race, Dr. Peter Tezla continues to study the information he gathered on the Wheel of Power while it was still in his possession. However, he finds that Gelorum has returned now fully repaired with her army of Racing Drones having been upgraded and she is attempting to get the Wheel of Power again. He enters Highway 35 himself and attempts to stop Gelorum but fails and is greatly injured just as the Drones finally capture the Wheel. He manages to escape back to Earth and the Drones begin to turn Hot Wheels City into their new headquarters.
Meanwhile, in present-day California, former World Race drivers Vert Wheeler, Kurt Wylde, Taro Kitano, and Mark Wylde have now become members of two street racing teams: the Teku and the Metal Maniacs. During a grudge race on the coastline, Dr. Tezla's robotic assistant Gig appears, telling the World Race drivers that Dr. Tezla needs their help. Arriving at Dr. Tezla’s old base from the World Race, with some of their new teammates for extra help, they find it partially destroyed (presumably by the Racing Drones). When former Dune Ratz driver Brian Kadeem appears and takes them to the AcceleDrome, Dr. Tezla's new base, which consists of a massive track that loops around the inside of a large cavern hidden inside a mesa, Dr. Tezla appears on a large hologram in the middle of the cavern and tells them what he has found: new Acceleron knowledge from the Wheel of Power's research. Now that the Wheel has been taken from its spire in Highway 35, the Drones have control over the dimension’s portals. However, the Wheel was not the final part of the Accelerons' test.
Each of the symbols on the Wheel represents one of the Racing Realms, inter-dimensional worlds that have tracks that go through a single element in the hardest way possible for driving conditions (there are only fifteen Realms in the series, twelve that were actually shown and three that the Drones had already completed before the beginning of the series, but only the AcceleChargers from those three are shown). The Drones have been entering the Realms, and each time a Realm is finished, the first driver to reach the end of the track obtains an AcceleCharger, which gives a car the ability to drive through any condition that relates to the realm that it was won in. Discovering that the Wheel hologram is always in sync with the actual Wheel, it functions as an inter-dimensional portal to the Racing Realms (it should also be noted that the Drones have their own Wheel hologram portal as well, due to them capturing the actual Wheel). The drivers enter the realms to try to stop the Drones, but find it harder than the World Race because the Wheel itself decides when to open the Realms and that each Realm only stays open for one hour with no possible way out.
As the race to stop the Drones rages on, drivers are trapped in the Realms, grudges between the teams grow, and a new enemy force known as the Silencerz appears, attempting to defeat both the Drones and the humans with the same Wheel hologram technology that the other drivers currently share. As the race to stop the Drones reaches its end, the secret of the Acceleron's test comes to unveil itself: the Racing Realms were designed to create "the perfect driver" and the AcceleChargers were created as keys to having the Wheel of Power open a portal to the Acceleron home world.